abnormal olfactory epithelium morphology, abnormal olfactory sensory neuron morphology, abnormal palate morphology, abnormal pancreatic beta cell differentiation, abnormal rostral-caudal axis patterning, abnormal sacral vertebrae morphology, abnormal sternocostal joint morphology, abnormal stomach epithelium morphology, abnormal tail morphology, abnormal vertebral column morphology, absent amacrine cells, absent retina inner plexiform layer, absent sacral vertebrae, absent tail, absent ureter, absent ureteric bud, asymmetric rib joints, cardiovascular system phenotype, caudal body truncation, cervical vertebral transformation, decreased caudal vertebrae number, decreased pancreatic acinar cell number, decreased pancreatic beta cell mass, decreased retina photoreceptor cell number, delayed optic fissure closure, homeostasis/metabolism phenotype, impaired branching involved in ureteric bud morphogenesis, increased apoptosis, increased cardiac stroke volume, increased lumbar vertebrae number, increased neuronal precursor cell number, increased pancreatic alpha cell mass, increased pancreatic islet number, increased presacral vertebrae number, increased retina ganglion cell number, increased thoracic vertebrae number, limbs/digits/tail phenotype, lumbar vertebral transformation, muscle phenotype, neonatal lethality, complete penetrance, pancreatic islet hyperplasia, preweaning lethality, complete penetrance, respiratory system phenotype, short tail, small kidney, small pancreatic islets, thoracic vertebral transformation, vertebral transformation